1. 《古DNA与人类历史》，本研合上课程。

2. 《分子考古学实践》，本研合上课程。

1. 国家自然科学基金面上项目-黄河中下游全新世以来古DNA视角的人群迁徙及其环境动因和对文明进程的影响（2025—2028）项目负责人，在研。

2. 国家优秀青年科学基金项目（海外）- 全新世东亚古代人群基因组学 （2023—2026）项目负责人，在研。

3. 全国考古人才振兴计划项目-三代都邑的墓地人群结构与血缘组织关系研究 （2024—2029） 项目第二负责人，在研。

英文文章(†共同一作; *通讯作者)

1. Zhang F., Gao S., Zhao X., Wu Y., Zhang J., Li J., Zhang J., Ning C.，Yan S., Wei D., Cui Y^*. 2025. Bronze and Iron Age genomes reveal the integration of diverse ancestries in the Tarim Basin. Current Biology, https://doi.org/10.1016/j.cub.2025.06.054

2. Wang J.^*, Yan S.^*, Li Z., Zan J., Zhao Y., Zhao J., Chen K., Wang X., Ji T., Zhang C., Yang T., Zhang T., Qiao R., Guo M., Rao Z., Zhang J., Wang G., Ran Z., Duan C., Zhang F., Song Y., Wu X., Ruth M., Sun B.^*, Pang Y.^*, Huang Y.^*, Zhang H.^*, Ning C.^* 2025. Ancient DNA reveals a two-clanned matrilineal community in Neolithic China. Nature 643, 1304–1311. https://doi.org/10.1038/s41586-025-09103-x

3. Wang J., Zhang T., Jia X., Zhang Z., He J., Ning C.，Wang J., Yang T., Wang G., Pang Y.^*, Huang Y.^* 2025. Genomic profiling of a six-generation patrilineal family of the Ming-Qing dynasties in China. iScience, 28, Issue 7, 112968.

4. Wang L., Duan C., Ning C^* 2025. Genetic insights into Ancient Kinship and Human History: Methods, Applications, and implications. Nature Anthropology, 3, 10009. https://doi.org/10.70322/natanthropol.2025.10009

5. Zhang J., Wang Y., Zhang N., Li J., Qu Y., Zhu C., Zhang F., Cai D.^*, Ning C.^* 2025. Unraveling the origins of the sogdians: Evidence of genetic admixture between ancient central and East Asians. Journal of Archaeological Science: Reports, 61,104957. https://doi.org/10.1016/j.jasrep.2024.104957.

6. Zhang F.^†, Chen Y.^†, Nie Z.Z.^†, Zhang R.J., Duan C., Wu D., Wang Y.Q.^*, Ning C.^* 2024. Complex genetic landscape revealed by a population in the eastern Tianshan Mountains of Xinjiang between the 8th and 11th centuries CE. International Journal of Osteoarcheology 34(4), e3300. https://doi.org/10.1002/oa.330010

7. Pei X.C., Cao P., Yang Y.D., Liew C.M., Zhang C., Qin L., Deng Z.H., Zhu S.Z., Chen Y., Zhao H., Ning C. Hudson M., Zhang H. 2024. Bone-artefact production in late Neolithic central China: evidence from Pingliangtai. Antiquity, 98 (399), 688-708. https://doi.org/10.15184/aqy.2024.56.

8. Zhang F., Zhang X.L., Bai B., Hu C.G., Duan C., Yuan H.B., Zhang R.J., Ma P.C., Zhou B.S., Ning C.^*. Ancient genomes provide insights into the genetic history in the historical era of southwest China. 2024. Archaeological and Anthropology Sciences. 15, 194. https://doi.org/10.1007/s12520-023-01899-x.

9. Zhang F., Liu Y., Ning C., Zhang J., Ma P., Zhang R., Yun Z.R., Duan C., Cai D.W., Yuan H.B.^* 2023. Ancient genomes reveal the origin and kinship burial patterns of human remains during the 11th to 13th centuries in northern China. International Journal of Osteoarchaeology, 34(1), e3274. https://doi.org/10.1002/oa.3274

10. Zhang F., Zheng L., Lv H., Duan C., Ma P., Liu Z., Zhou K., Tang F., Ning C., Zhang Y.,2024. Ancient Genome Analyses Shed Light on the Origin and Kinship among Humans of a Cliff Tomb from Southwestern China. Journal of Archaeological Science, Reports 53, doi: https://doi.org/10.1016/j.jasrep.2023.104333.

11. Zhang N., Liang Q., Shao X., Guo Y., Wang Y., Wang X., Zhang Wei., Ning C., Cai D., 2023. Ancient Cattle DNA Provides Novel Insight into the Subsistence Mode Transition from the Late Neolithic to Bronze Age in the Nenjiang River Basin. Journal of Archaeological Science, Reports 51, doi: https://doi.org/10.1016/j.jasrep.2023.104136.

12. Cai D., Zheng Y., Bao Q., Hu X., Chen W., Zhang F., Cao J., Ning C. 2023. Ancient DNA Sheds Light on the Origin and Migration Patterns of the Xianbei Confederation. Archaeological and Anthropological Sciences 15, no. 12, doi: https://doi.org/10.1007/s12520-023-01899-x.

13. Chen L., Ning C., Ma L., Wang M., Zhao Y., Li S., Zhao Z., 2023. Harnessing the Potential of Postbiotics Derived from Bacillus Coagulans: A Promising Avenue for Obesity Management Via the Gut-Liver Axis. Food Bioscience 56, doi: https://doi.org/10.1016/j.fbio.2023.103108.

14. Liang Q., Chen Q., Zhang N., Zhang W., Ning C., Cai D.,2023. Subsistence Strategies in the Late Neolithic and Bronze Age in Nenjiang River Basin: A Zooarchaeological and Stable Isotope Analysis of Faunal Remains at Honghe Site, Northeast China. International Journal of Osteoarchaeology 33, no. 2, doi: https://doi.org/10.1002/oa.3212.

15. Zhang H., Pei X., Li W., Zhang F., Liang F., Cao Y., Liu C., Wei X.*, Ning C.*, 2023. Millet Diffusion and Subsistence Diversification in Late Neolithic Central China: Perspective from Archaeological, Archaeobotanical, Stable Isotopic and Genetic Evidences. Annals of Archaeology 5, no. 1, doi: 10.22259/2639-3662.0501003.

16. Zhang F.†, Ning C.†,*, Scott A.†, Qiaomei Fu, Rasmus Bjørn, Wenying Li, Wei D., Wang W.3, Fan L., Abuduresule I., Hu X., Ruan Q., Niyazi A., Dong G., Cao P., Liu F., Dai Q., Feng X., Yang R., Tang Z., Ma P., Li C., Gao S., Xu Y., Wu S., Wen S., Zhu Hong, Zhou Hui, Robbeets M., Kumar V., Johannes Krause J.*, Warinner C.*, Jeong C.*, Cui Y.*, 2021. The genome origin of the Bronze Age Tarim mummies. Nature 1-6 doi:10.1038/s41586-021-04052-7.

17. Robbeets M.*, Bouckaert R., Conte M., Savelyev A, Li T., Deog A., Shinoda K., Kim J., Cui Y.Q., Kawashima T., Kim G., Uchiyama J., Oskolskaya S., Dolinska J., Deng B.C., Bjørn R., Yamano K.Y., Seguchi N., Tomita H., Takamiya H., Kanzawa H., Oota H., Ishida H., Kimura R., Sato T., Rhee S., Ahn K.D., Gruntov I., Bentley J., Fernandes R., Roberts P., Bausch I., Gilaizeau L., Yoneda M., Kugai M., Bianco R., Zhang F., Burri M., Himmel M., Hudson M.*, Ning C.*, 2021. Triangulation supports agricultural spread of the Transeurasian languages. Nature doi: 10.1038/s41586-021-04108-8

18. Wu X.Y., Ning C., Key F., Valtuena A., Lankapalli A. K., Gao S.Z., Yang X., Zhang F., Liu L.L., Nie Z.Z., Ma J., Krause J.*, Herbig A. *, Cui Y.Q. *. 2021. A 3,000-year-old, basal S. enterica lineage from Bronze Age Xinjiang suggests spread along the Proto-Silk Road. PLOS Pathogens 17, e1009886

19. Ning, C., Zhang, F., Cao, Y., Qin, L., Hudson, M.J., Gao, S., Ma, P., Li, W., Zhu, S., Li, C., Li, T., Xu, Y., Li, C., Robbeets, M., Zhang, H., Cui, Y., 2021. Ancient genome analyses shed light on kinship organization and mating practice of Late Neolithic society in China, iScience, doi: https://doi.org/10.1016/j.isci.2021.103352.

20. Zhao Z.J., Ning C.*, Chen L., Zhao Y.J., Yang G., Wang C., Chen N.S., Zhang Z.Y., Li S.Y.*, 2021 Impact of manufacture processes and geographical regions on the microbial profile of traditional Chinese cheeses. Food Research International.148:110600

21. Ning C., Zheng H.X., Zhang F., Wu S.H., Li C.X., Xu Y., Gao S.Z., Jin L., Yinqiu Cui Y.Q., 2021. Ancient Mitochondrial Genomes Reveal Extensive Genetic Influence of the Steppe Pastoralists in Western Xinjiang. Front. Genet. https://doi.org/10.3389/fgene.2021.740167

22. Ning C.†, Fernandes D.†, Changmai P.†, Flegontova O.†, Yüncü E.†, Maier B., Altınışık E., Kassian L., Krause J., Lalueza C., Manica A., Potter B., Robbeets M., Sirak K., Siska V., Vajda E.J., Vyazov L.A., Wang K., Wang L.X., Wu X.Y., Xiao X.M., Zhang F., Reich D., Schiffels S., Pinhasi R.*, Cui Y.Q.*, Flegontov P.*, 2020. The genomic formation of First American ancestors in East and Northeast Asia. bioRxiv. doi: https://doi.org/10.1101/2020.10.12.336628.

23. Ning C., Li T., Wang K., Zhang F., Li T., Wu X., Gao S., Zhang Q., Zhang H., Hudson M., Dong G., Wu S., Fang Y., Liu C. Feng C., Li W., Han T., Li R., Wei J., Zhu Y., Zhou Y., Wang C., Fan S., Xiong Z., Sun Z., Ye M., Sun L., Wu X., Liang F., Cao Y., Wei X., Zhu Hong, Zhou Hui, Krause J., Robbeets M., Jeong C., Cui Y., 2020. Ancient genomes from northern China suggest links between subsistence changes and human migration Nat. Commun. 11, 2700.

24. Yang M., Fan X., Sun B., Chen C., Lang J., Ko Y., Tsang C., Chiu H., Wang T., Bao Q., Wu X., hajdinjak M., Ko A., Ding M., Cao P., Yang R., Liu F., Nickel B., Dai Q., Feng X., Zhang L., Sun C., Ning C., Zeng W., Zhao Y., Zhang M., Gao X., Cui Y., Reich D., Stonking M., Fu Q., 2020. Ancient DNA indicates human population shifts and admixture in northern and southern China. Science. doi:10.1126/science.aba0909.

25. Ning C., Wang, C., Gao, S., Yang, Y., Zhang, X., Wu, X., Zhang, F., Nie, Z., Tang, Y., Robbeets, M., Ma, J., Krause, J., Cui, Y., 2019. Ancient Genomes Reveal Yamnaya- Related Ancestry and a Potential Source of Indo-European Speakers in Iron Age Tianshan. Curr. Biol. CB 29, 2526-2532.e4. https://doi.org/10.1016/j.cub.2019.06.044.

26. Li, T., Ning C., Zhushchikhovskaya, I.S., Hudson, M.J., Robbeets, M., 2020. Millet agriculture dispersed from Northeast China to the Russian Far East: Integrating archaeology, genetics, and linguistics. Archaeol. Res. Asia 22, 100177. https://doi.org/10.1016/j.ara.2020.100177.

27. Ning C., Gao, S., Deng, B., Zheng, H., Wei, D., Lv, H., Li, H., Song, L., Wu, Y., Zhou, H., Cui, Y., 2016a. Ancient mitochondrial genome reveals trace of prehistoric migration in the east Pamir by pastoralists. J. Hum. Genet. 61, 103–108. https://doi.org/10.1038/jhg.2015.128.

28. Ning C., Yan, S., Hu, K., Cui, Y.-Q., Jin, L., 2016b. Refined phylogenetic structure of an abundant East Asian Y-chromosomal haplogroup O*-M134. Eur. J. Hum. Genet. 24, 307–309. https://doi.org/10.1038/ejhg.2015.183.

29. Li C†., Ning C†., Hagelberg, E., Li, H., Zhao, Y., Li, W., Abuduresule, I., Zhu, H., Zhou, H., 2015. Analysis of ancient human mitochondrial DNA from the Xiaohe cemetery: insights into prehistoric population movements in the Tarim Basin, China. BMC Genet. 16, 78. https://doi.org/10.1186/s12863-015-0237-5.

30. Cui, Y., Zhang, F., Ma, P., Fan, Z., Ning C., Zhang, Q., Zhang, W., Wang, Robbeets, M., 2020. Bioarchaeological perspective on the expansion of Transeurasian languages in Amur River Basin. Evol. Hum. Sci.

31. Juno U.*, J. Christopher, G., Alexander, S., Ning C.*, 2020. Populations Dynamics in Northern Eurasian Forests: A Long- term Perspective from Northeast Asia. Evol. Hum. Sci.

32. Wu, X., Ding, B., Zhang, B., Feng, J., Wang, Y., Ning C., Wu, H., Zhang, F., Zhang, Qun, Li, N., Zhang, Z., Sun, X., Zhang, Quanchao, Li, W., Liu, B., Cui, Y., Gong, L., 2019a. Phylogenetic and population structural inference from genomic ancestry maintained in present-day common wheat Chinese landraces. Plant J. Cell Mol. Biol. 99, 201–215. https://doi.org/10.1111/tpj.14421.

33. Wu, X., Ning C., Bao, Q., Gao, S., Zhang, F., Wu, S., Li, Tianjiao, Fan, L., Li, Tao, Yang, X., Cai, D., Cui, Y., 2019b. Mitochondrial Genome of an 8,400-Year-Old Individual from Northern China Reveals a Novel Subclade Under C5d. Hum. Biol. 91, 21–30. https://doi.org/10.13110/humanbiology.91.1.04.

34. Cui Y., Li H., Ning C., Zhang Y., Chen L., Zhao, X., Hagelberg E., Zhou, H., 2013. Y Chromosome analysis of prehistoric human populations in the West Liao River Valley, Northeast China. BMC Evol. Biol. 13, 216. https://doi.org/10.1186/1471-2148-13-216.

35. Cui Y., Song, L., Wei, D., Pang, Y., Wang, N., Ning C., Li, C., Feng, B., Tang, W., Li, H., Ren, Y., Zhang, C., Huang, Y., Hu, Y., Zhou, H., 2015. Identification of kinship and occupant status in Mongolian noble burials of the Yuan Dynasty through a multidisciplinary approach. Philos. Trans. R. Soc. Lond. B. Biol. Sci. 370, 20130378. https://doi.org/10.1098/rstb.2013.0378.

36. Han L., Zhu, S., Ning C., Cai, D., Wang, K., Chen, Q., Hu, S., Yang, J., Shao, J., Zhu, H., Zhou, H., 2014. Ancient DNA provides new insight into the maternal lineages and domestication of Chinese donkeys. BMC Evol. Biol. 14, 246. https://doi.org/10.1186/s12862-014-0246-4.

37. Hu K., Yan, S., Liu, K., Ning C., Wei, L.-H., Li, S.-L., Song, B., Yu, G., Chen, F., Liu, L.-J., Zhao, Z.-P., Wang, C.-C., Yang, Y.-J., Qin, Z.-D., Tan, J.-Z., Xue, F.-Z., Li, H., Kang, L.-L., Jin, L., 2015. The dichotomy structure of Y chromosome Haplogroup N. ArXiv150406463 Q-Bio.

中文文章

1. 崔银秋，方燕明，宁超，李天娇. 河南禹州瓦店遗址龙山时期古代居民DNA分析. 华夏考古. 2014 (4): 41-46.

2. 杨翊研，宁超，杨百全，崔银秋，李春香. 采用STR、SNP和X-STR进行胚胎组织亲权鉴定1例. 中国法医学杂志. 2016 (31-4).

3. 王尹辰，马鹏程，张帆，宁超，高诗珠，马健，崔银秋. 早期铁器时代新疆东天山地区与欧亚草原的基因交流. 西域研究. 2020 (3): 106-172.

学术专著

1. Brandley D., Liu L., Ning C., Martello R. 2025. Sino-Tibetan archaeolinguistics. In the Oxford Handbook of Archaeology and Language (eds. Robbeets M., Hudson M.)

2. Robbeets M., Hudson M., Ning C. 2025. Transeurasian archaeolinguistics. In the Oxford Handbook of Archaeology and Language (eds. Robbeets M., Hudson M.)

3. Jeong C., Wang C., Ning C., 2020. Transeurasian unity from a population-genetic perspective. In the Oxford Guide to the Transeurasian Languages (eds. Robbeets M., Savelyev A.)

专利

Cui Y., Zhao H., Zhang X., Ning C.. Animal species identification kit for degraded samples. ZL 2014 1 0465741.7. 2016.

参加会议(节选)

Ning C.. Archaeogenetic evidence for Neolithic migration and Bronze Age interation in North and East Asia (德国)，2021 (大会报告-网上)

Ning C.. Population genetic perspective on the Neolithic populations in Northeast Asia. Workshop for ICHL24, Australian National University, Australia (澳大利亚), 2019 (大会报告).

Ning C., Population genetic history in East Asia. Fourth Annual DFG Center Symposium: Ancient Connections in Eurasia, Germany (德国), 2018 (大会报告).

Ning C., Wang C. Genomic insight into the peopling of Northeast Asia. Society for East Asian Archaeology, Nanjing (南京) 2018 (大会报告).

Ning C., Genomic insight into the peopling of northeast China. International Society for Biomolecular Archaeology, Germany (德国) 2018 (大会报告).

Ning C., Zhang H. Millet spread and complex subsistence in Late Neolithic of Northeast China: Evidence from archaeology and genetics. Transeurasian millets and beans, language and genes, Germany (德国) 2019 (会议组织者及大会报告).

Ning C., Genomic perspective on the peopling of Transeurasian region. Sainsbury Institute, University of East Anglia, UK (英国) 2018 (大会报告).